BENMOUNA, Zahia, University of Sciences and Technology of Oran Mohamed Boudiaf, USTOMB, Faculty of Natural and Life Sciences, Department of Living and Environment, PO Box 1505, El Mnaouar, Oran 31000; University of Science and Technology Houari Boumediene, USTHB, Faculty of Biological Sciences, Laboratory of Research in Arid Areas, PO Box 32, El Alia, Bab Ezzouar, Algiers 16111, Algeria